Cambridge Entomological Club, 1874
PSYCHE

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Article beginning on page 39.
Psyche 9:39-40, 1900.

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PSYCH EL
A NEW BLEPIIAROGER-ID.
HY VEIZNOX I.. KELLOGG, STANFORD UNIVERSITY, CALIF. The family of Blepharoce~~iilae, as at
present known, includes about fifteen
species of nematocerous Diptera, of
unusual interest to entomologists. This
interest is caused by the rarity of speci- niciis in collections, by the unique
structural condition of the larva, by
the strange, although as yet imperfectly known, lilt history and habits, and
finally by the peculiar accessory vena-
tion :md suggestive structural character of the mout!iparts and *corupound eyes
of the imagines.
In the summer of 1895, Mr. R. W.
Doiine, at that time collecting- for me in the vicinity of this University (Stan-
Ford), took two females and a male of
a Blepharocerid species. I have only
recently given these specimens any
attention other than the rather unkind
one of removing the heads of two (the
male iind one female) in order to study
their mouthparts. A recent examina-
tion of the specimens reveals the fact
that they are reprtsentiitives of an unde- scribed species which may be assigned
to Low's gonus Liponcura (Stett. Ent.
Zeit. 1344, vcl. v, p. 118).
... .
.-
*See Kellogg, Notes on the Lie-history and Structure of Blepliarocera capitata Low; Eut. News, 1900, vol. X, pp. 305-318.
The new species may be described as
follows :
Liponem dmwi 11. sp.
Female; length
6 mm., length of wing 74 mm.; very pills brown, almost clavey ; iintennne 14-seg- merited and rising from a prominence which Fig. i. Mouthparts of L&weura doanei
11. sp.
might be construed to be a basal antenna1 segment; eyes broadly separated, and with no indications of bisection, the facets being Pswhe 9 039-41 (pre.1903) http //psyche aitclub ora/9/tl.OOJ9 html



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Ill PS VCIfE. [.April
all of tlic same six: the nioiithparts (fig. I) long (distinctly longer than in Blepharocera, for example) and composed of long slender flattened well chitinized mandibles with ilie apical two-fifths of the inner margin finely serrate, and inserted farther back on the head thnn the other mouthparts; of maxillae conxisting <if slender flattened tapering blade- like terminal lobes and long five-segmented palpi; of elongate simple labium consist- ing- of basal portion suid one pair of free terminal lobes; and of slender elongate Qat- tened labrrii"!.epipharynx and hypopharynx; wings wit11 venation (fig. 2) showing the fol- lowingcharsicters (given first in the nomen- clature used by Combtock and in following parentheses in the nomenciaitire used by Osten Sixken in the latcst revisional paper Di~sal rudiment present (;uixiiiav vein n ing or with only a basal rudiment press
Wings clear, with strong iride scent re
tions. The legs are long, the three paii about equal length, the hind tibiae beari single terminal spur, the other tibiae w out spurs. The ti~rsal claws are large, t and strong with curved pointed tip, !In
pectinate except at the tip. The exte
genitalin consist of a pair of flattened airgular dorsal plates.
Male. The inale is slightly smaller t
the female.
As the head of the on11 I
specimen was destroyed in the dissectio
the mouthparts before any exainiiiatiot
the specimen was made, it csuinot be
whether the eyes are like those of the fel or not; mouthparts like those of the fern except that the inandi
are wholly lacking. Wi
and legs like female. '
conspicuons parts of
external genitalia an
pair of large artict~li
claspers, a broad ven
plate, and a smaller do
plate.
Fig. 2. Venation of Liponevra doanez 11. sp. Desciibed fiom I
of the Blepharoceridae) ; vein M3 independ- ent, i, e. without connection with M or any other principiil vein (an incomplete vein running into the posterior margin between veins 4 and 5); veins Rn and R, coalesced to the margin (vein 2 simple, iinbranched) ; a medio-cubital cross vein (a cross-vein between veins 4 and 5): veins Ra+, and
R4+s separating at the origin of the radio- medial cross vein (the sub-marginal cell sessile); the radial sector springing from R by two roots (the cross vein between
veins I and 2 Y-shaped, that is the anterior half of it divided, enclosing a small triangu- lar cell) ; sub-costa wanting or with only a . --
Osten Sackeii C. R.
Contributions to the study of the
females and one male, taken by R.
Doane, July 24, 1895, on the banks
a small stream in the Santa Cruz IV
at Congress Springs, Santa Cl
County, California.
The new species can readily
grouped with the four other kno
species assigned to the genus Liponei
in Osten Sacken's* paper. The n
form has in common with the otl
four species an incomplete vein runni
into the posterior margin of the w;
between veins 4 and 5, a simple (i
Osten Sackeii C. R. lor. cit




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branched) second longitudinal vein,
mcl eyes separated by a broad front.
Within the genus, the new form agrees
with bilobata Low and yosemite 0. S.
in possessing a cross vein between
veins 4 and 5, and finally resembles
bilobata in having the sub-marginal
cell sessile. But it differs from bilobata (as from till other Blepharoceridae) in
having the radial sector springing from
two roots (the base of the second longi- tudiliiil vein forked) so that a small tri- angular cell is formed behind Rl (first
longit~~dinal vein).
The only other Blepharoceridae so
far known from the Pacific Coast arc
Blefharocera ancilla 0. S. (Cali-
fornia) and Li-eura yosemite 0. S.
(Yosemite Canon, California), from
both of which the new species differs
sharply in the character of the eyes and venation.
Unfortunately I have not been able
vet to find the immature stages of the
new species, so can add nothing to our
incomplete knowledge of the interest-
ug life-history of the members of the
family.
One of the moot points regarding
the biology of the Blepharoceridae is
PROCEEDINGS OF THE CLUB.
12 JANUARY, ~yoo. The 211th regular and
23d annual meeting (since incorporation) was held at 156 Brattle St., Mr. W. L. W. Field in the chair
Repork from the several officers were
received and the following officers elected for the ensuing year:-President, J. W.
Folsom ; secretary, Roland Hayward ; treas- urer, Samuel Henshaw; librarian, Samuel H. Scudder; members at large of the executive committee, A. S. Hewins and A. P. Morse. that of the dimorphisni of the female.
I have elsewhere * referred to Fritz
Muller's statement that there are two
kinds of females of Paltostoma towen-
tilt-m (Brazil), one kind possessing
mandibles and being blood-sucking,
the other kind having no mandibles
and being nectar-sucking. Osten
Sacken deems the evidence of dimor-
phism insufficient. Of twenty-three
females of fileflurocera capitata
LOW taken by me at Ithaca, N. Y.,
no one was without mandibles, nor
was there any other difference appar-
ent. Most of these specimens were
taken just as they were issuing from
the pupal skins on various days, in
various parts of the stream, so the crit- icism that one kind of female might
possess habits rendering it more likely
to be caught than the other, will not
hold in this instance. It seems to me
probable that there is no dimorphism
of the females of Blepharocera ca'pt-
tata Low. In the case of the new
Cnlifornian species I can only say that
both females (the only ones so far
taken) agree in possessing mandibles,
and in all other characters.
The address" of the retiring president,
A. G. Mayer, on the mating instinct in
moths was next read. (See Psyche for Feb- ruary.) Much discussion followed, in'which all present participated.
Mr. W. L. W. Field spoke of a cocoon of
Samia cyntJtia which he had seen contain- ing two pupae. In outward appearance,
however, it showed no apparent difference from a normal cocoon of that species.




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