Article beginning on page 207.
Psyche 7:207-211, 1894.

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PSYCHE.
NEW NORTH AMERICAN ODONATA.
BY ALBERT P. MORSE, WELLESI,EY, MARS.
Calvert, in his catalogue of the
Odonata of Philadelphia, lists the
following eight species of EnuZlagma
from the northeastern States : aspw-
sum, civile, divtzgans, dzwztĺ´~~z exsu- lam, haye&, poZlut1tn2, signaturn.
Other forms reported by various
authors from this district are : an-
nexum, ebrimz, tru~iatum. To these
Calvert (Ent. news, Oct. 1894) adds
douhledayi; and in the following
pages two more are added, making
fourteen species of the genus to be
found in New England. One of the
latter (und two additional new species
described here) is found in Nevada
also. A note is added on the diag-
nostic rlifTercnccs of asjersum and
traviatmn. Descriptions are also pre-
sented of an Ophiogonzfhus from the
eastern States, a Tei!ra,qonettria from
Massachusetts and a case of variation
in venalion in ~ryt/;?-omina con&um.
In consideration of the fact that thu
species of EnaZlagma approach each
other so closely that sketches of ana-
tomical details are almost imperatively
demanded, the author intends to pre-
sent in a subsequent paper diagnostic
drawings of all the New England
species and of as many other North
American forms as it may be possible
to obtain.
Being a firm believer in the desira-
bility of assembling the types of closely allied species in order to facilitate
comparative study, the writer proposes
to ultimately present the types here
described to the Hagcn collcction in
the Museum of Comparative Zoology
at Cambridge, Mass.
Enallagma rninusculum, sp. nov.
Abd. 20 mm., hind wing, 14-15 inm.
Cuneiform post-ocular spots small, more
or less rounded. Posterior lobe of prono- turn black, unspotted,margined with yellow; anterior lobe with pale transverse band. Thorax with the following black: a wide
mid-dorsal stripe, widest in front; a wide hu~nural (wilier than pale ante-lnimcrai) stripe, widest below, expanded on and just behind its crossing of the humeral suture. Abdomen black as follows: dorsum of I,
divided by an apical spot of blue or purple; posterior half of 2, third of 3, two-fifths of 4, half of 5, three-fifths of 6, four-fifths of 7, and all of 10; 8 find g blue.
Superior appendages half as long as 10,
in profile broader than long, equalling in depth (on expanded portion) one-half tliat of 10, very broadly liifid, the posterior margin shallowly excavate, the lower
brimuh projecting a little beyond the upper, much expanded, laminate, directed down-
ward and inward; the upper branch one-
third as broad, directed slightly upward, and bent inward claw-like. Inferior appen- dages equal in length to the superior, broad at base, narrowed and tapering in the




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208 PSYCHE. [March i895
middle portion, the upper margin of which runs directly backward, bent upward and
slightly inward at the tip.
7 3, Sherborn, Mass., Jul? 16.
Collected
by A. P. Morse.
Enallagma calverti, sp. nov.
Abd. 23-25 mm., hind wing 17-10.5 inm.
Prothorax greenish black, the following
pale (bluish) : sides; a transverse line on anterior lobe; the hind margin and a
cuneiform spot on each side of posterior lobe. Thorax with a rather narrow mid-
dorsal stripe (sometimes divided by a mere line of blue, most distinct anteriorly) and a very narrow humeral stripe, wider in
front, especially at the suture, and a spot on second lateral suture, black. A wide
ante-humet-al stripe, equal to or wider than the mid-dorsal black stripe, blue, Abdo- men blue, the following black: a spot on base of I ; a transverse lunule (convex side forward, doubly concave behind) near apex and a narrow marginal band on 2; an
apical spot connected with marginal band on 3 and 4; apical third of 5, two-thirds of 6, five-sixths of 7, and all of 10.
Superior appendages short, one-fourth to one-third as long as 10, blunt, with the apex directed downward and slightly
notched in profile; the upper limb thick and rolled inward, the lower limb thin,
rolled inward and upward, appearing like a small, rounded, inwardly projecting shelf on the lower edge of the apex of the
appendage. In profile the upper apical
angle is very obtusely rounded, the lower slightly notched. Inferior appendages
longer, two-thirds as long as 10, rather lender, tapering, slightly curved upward, directed upward and backward, the"Tower
margin convex throughout.
g 3, Franktown, Nev., June (coll.
by S, W. Denton).
To this species I also refer I 3,
Wellesley, Mass., May 12 ; I if, prob-
ably from Wellesley, presented by S. F.
Denton ; and I $, eastern U. S.,
exact locality uncertain, the only
difference apparent being that the
inferior appendages in profile seem to
be somewhat more slender and less
strongly convex below.
I take pleasure in naming this
species for Mr. P. P. Calvert, whose
" Introduction" to this group of our
fauna fills a long-felt want and will do much to promote the study of these
interesting insects.
Enallagma carunculatum, sp. nov,
Abd., 22.5-27; hind wing, 17-20 mm.
Prothorax with the anterior lobe trans-
versely lineate with pale; posterior lobe with pale margins and cuneiform lateral
spots; the latter arc small, narrow, or even may be wanting. Thorax with the mid-
carina black or lineate with pale; mid-
dorsal dark stripe broad; humci'iil stripe of moderate width, variable, widest just behind suture; ante-humeral pale stripe
moderate to narrow, varying from ncwly
twice as wide to only two-thirds as wide as the humeral dark stripe. Abdomen black as follows: a quadrate spot on base of I; an apical orbicular spot broadly connected to posterior marginal band on 2; apical
half to three-fifths of 3, 4, 5 with a longi- tudinal band, pointed anteriorly, widened before margin; two-thirds to three-fourths of 6: all of 7 except a narrow basal ring; all of 10, widest on basal and apical mar- gins; 8 and 9 blue.
In profile the superior appendage, inclnd- ing the projecting tubercle, is half to two- thirds as long as 10, the inferior appendage shorter, equal to sup. app. without tubercle, stout, directed moderately upward, the upper margin concave, the lower strongly convex. Superior appendage as broad at base as the



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March i8~s.l PSYCHE.
inf. app., about as broad ELS long, the sides equal, nearly straight, the upper slightly convex. the lower sli:litly concave, the apical margin concave, bearing a large,
yellow or brown tubercle, rounded apically. two-thirds as long as the appendage; the groove separating it from the end of the appenci~ige is more distinct in he dorsal portion, appearing' like a slight notch, recalling E,. chile. Seen from above the superior appendages are soinewhat divari- cated, cylindrical, terminating apically in a rounded right-angle in the mid-line of the appendage; the tubercle projecting
beyond, narrower, longer than broad,
rounded apically: the inner margin of the appendage forms a minute tooth, vci-J
slightly recurved, at the obtuse inner apical angle. Seen from behind the dorsal and
ventral edges of the superior appendage
are seen to be rolled inward, the tubercle occupying this inner channel and (in dry specimens) slightly rolled, also, at its tip. 12 (^ (I headless, I teneral), Franktown: Nev., June. Collected by S. W. Denton.
Enallagma clausum, sp. uov.
Abdomen 25-28; hind wing, 19-21 mm.
Prothorax with the anterior lobe trans-
versely lineate with pale; posterior lobe greenish black, with pale margins and
triangular or cuneiform lateral spots.
Thorax with the mid-carina black or
ineate with pale; mid dorsal dark stripe rather narrow; humeral stripe very narrow, widest and more or less broken at the
suture. Pale ante-humcral stripe at least two-thirds as wide as mid-dorsal dark
stripe. Abdomen black as follows : a quad- rate spot on base of I ; a large apical orbi- cular spot, pointed anteriorly and broadly conected to narrow hand on apical margin of 2; apical two-fifths to two-thirds of 3, 4, 5 with a longitudinal stripe narrowed submarginally (narrowest and pointed
anteriorly on 3) ; apical two-thirds of 6, four-fifths of 7, and all of 10, on the latter broadened marginally, especially on basal margin ; 8 and 9 blue.
In profile the inferior appendage is the longer, two-thii-ds as long as 10, stout, directed nearly straight backward, the
lower margin strongly convex, the upper
strongly concave, the apex stout, upLurned, about on a level with the upper part of the broadened base; the whole appendage very similar in shape to a cut's claw, but less acute at tip. Superior appendiige two-thirds as long as infprior, directed downward,
the lower margin slightly descending
toward sipex, the upper margin convex
hasally, slightly sinunte at apex, some- what recalling t11a.t of 23. divagans but less hollowed near apex and less convex toward base. In dorsal view the hind margin of
10 is deeply excised, with a deep depres- sion in the middle between the dorsal
border and the superior appendages. Supe- rior appendages short, blunt, very broad, cowtiguous at base for nearly half their length, the line of separation often visible only with difficulty,-the contiguous por- tion with rugose inner margin terminating apically in a larger, slightly recurved tooth; beyond the tooth the appendage is suddenly narrowed on the inner margin for a third of its withh, the re-entrant angle nearly square, the distal inner margins of the pair of
appendages diverging moderately to the
rounded apices; the outer margin is convex basally, nearly straight apically.
6 8, Franktown, Nev., June.
Collected
by S. W. Denton.
Ophiogomphus aspersus, sp. nov.
Abd., 8, 30 mm.; 7, 30-32. Hind wing,
(T. 24; 9, 26-27. Hind fern., 7. Pter., 3.5, yellowish-brown, margined with black.
Wings very slightly flavescent at base. Face yellowish-green. Vertex black, posteriorly brown and green. Occiput yellowish-green ; back of head immediately behind eyes black above, on sides greenish-yellow, paler below. Occipital horns slender, brown. Thorax'




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PSYCHE,
[March 1895.
yellowish-green, marked with brown as
follows: a narrow mid-dorsal band, widest in the middle, narrowed at each end, in front barely wider than the mid-canna which may be partly pale, continuous behind with the brown of the ante-alary sinus; a very
narrow humeral stripe, widest above, where it is partly divided by a pale streak on the suture; and an equally nurow ante-htimeral stripe separated from the humeral by a
green band of equal width which is contiiui- ous above with the green of the dorsum,
thus separating the ante-humeral stripe from the sinus. Legs with the basal 8 of femora green (sometimes fuscous above), otherwise black. Abdomen dark In-own marked with
yellow as follows: & dorsum of I i1nd 2; basal half to two-thirds of 3-8 with a tri- ang~ilar spot, produced apically; 9 with :I quadrate or oi-bicular spot on basal third3 with an apical tdil; 10 with a lanceolate spot,
its apex reaching the hind margin;
ventral portions of sides of 8 and 9 and indications on the same portions of
preceding segments. si~nilar, more
suffused. 10 one-half as long as 9 in both sexes. Vulvar lamina + as long as 9, biiid for a little more than its apical half, the branches slightly divergent at first, then convergent, the tips contiguous, their apices rounded or with a minute tooth, the basal inner half of each branch somewhat thick- ened. Superior appendages of (^ as long as 10 am1 fs of 9. Seen from above the basal half is 8 as broad as long, the distal half smoothly tapering, with straight side>,
icuLe, sharp ; the inner margin of the basal half is deeply concave, the outer margin slightly so. Seen from the side the sup. app. is a little upturned at the tip, acute, almost acuminate; the dorsal margin
strongly sinuak, being slightly concave at the basal fourth, stron:Iy convex; at thc half, and distinctly concave on the distal third. Its lower margin is deeply excavate at the basal fourth, strongly convex and denticulated on the remaining portion; the distal <I of the outer ventral face of the appendage is convex in both longitudinal and transverse section and thickly beset with black denticles; the narrowed basal part is a little stouter than the corsespond- ing part of the inf. app. Inferior appendage $ as long as sup. app., the lower margin of the branch straight, slightly ascending, the uppermargin concave on the basal f,
straight and descending on the apical third, forming a tooth-like process at the two- thirds point and an acute apex,
2 & 3 Q, northeastern U. S. (probably
Wellesley, Muss., but if not, then Ohio, Ind., or Ill.); onc 9 received from S. F. Denton, the remainder From S. W. Denton. Tetragoneuria indistincta, sp. nov.
Very similar Lo 2'. cynoswa Say, but
somewhat Isn'gcr and differing from it as follows: a black 'r spot on front above; pubescence of thorax deep brown, instead of somewl~at hoary; wings slightly fi11110se and a little (2 inin,) longer, the bnal part of median space and basal antecubital cell of cacl~ series fuscuus; ptcrostigma slightly longer; abdomen longer (2 nun.), more
slender, especially toviiril apex, ninth segment nearly as Iongas the width of its posterior margin, 10 also distinctly longer proportionally than in cposura; superior appendages 2.7 nim. long, fusifor~n-cylindric, a little slenderer at base, their apices rounded (of cynosura 1.8 mm. long, and the apiccs sharply pointed).
Abdomen 29 mm. ; hind wing, 31. I 9,
Winchendon, Mass., July 2.
This species has probably been con-
fused with T. cynosura Say. In fact,
much confusion has existed and
perhaps still exists concerning the
01-ins known as cynoswa, semiaqnea,
and othcrs closely allied to them.
I regret to say that even the specimens
in the Hagen collection arranged under
these names are sadly mixed, the same




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March 1895.1 PSYCHE. 211
species occurring under two or three
names, and two or three species
occurring under one name. The
species referred to here as cynosura is
the one abundant in Massachusetts in
June, and presents two forms, as
follows: (I) that known as cynoszwa,
having the fuscous marking of the hind
wings restricted to the immediate base
( 9 ), or basal and anal parts (8 ) of
wings; 'Am\ (2) what has been called
semiapea, having the fuscous extend-
ing from the anal angle nearly to the
nodus, with a slight nodal spot which
is sometimes connecf.ed 1.0 the base;
specimens with markings intermediate
in character are not uncommon,
varying froin a mere f~~scuus bordering
of the venules of the anal area up to the form described here. These two forms
are doubtless one species, no structural differences being perceptible.
There is another species foiim-1 in the
southern Slates, having the fuscous of
a mole rcddish hue, and even wider
in extent, which presents differences in the abdominal appendages. This is
perhaps the tiue semiayuea.
Erythromma conditum Hng.
A series of 3 $, z 9, specimens of this
species collected by Mr. S. W. Denton,
probably at Wellesley, Mass., presents an in- terestiug case of variation in venation. It is the more worthy of attention for the
reason that the variation Ei/Sccts a character upon which analytical keys are often based. In one female the lower sector of the tri- angle arises distinctly before the basal post- costal nei-vule, in the fore wings about twice as fur as in the hind uings; in the other female it arises at Lhe pnstcostal nervule on all wings. In two males the sector arises just before it on the front wings and at it on the hind wings; in the other male at or al- most imperceptibly before on one hind wing, very slightly before on the other, and dis- tinctly before on thefront wings.
Enallagma traviatum Selys vs. E. aspersum Hag.
litinks (Cm Ent.. Mar. 1894) considers
Craviatitni. s1s "hardly more than a variety of a.-$ersum." From this opinion I must
dissent, regarding it entitled to full specific rank, a numberof males collected last summer showing marked differences when compared with aspersum. These cliITerences, in ad- dition to the characters indiciited in the original description of travi'aiuni (Syn. d. Agr.), me a5 fo1lo~-s : 1st: side view, trti-via- turn: Superior abdominal appendage with
the upper branch slender, nearly- equal
throughout, slightly cleciu-ved, the apex equally rounded, not especially dccnrvcd. Inferior appendage projxting one-thii-tl its length beyond the lower branch of the sup. app., the upper margin nearly straight, the lower margin deeply excavated, the exca- ration dividingtthe app. into a broad base and a slender tip, the base being about four times as broad and one-half as long as the tip, which is slender, equa.1, striught, and directed nearly straight backward. Side view, as*- sum: Sup. app. with upper branch rather
stout, its apex noticeably dccurved. Inf. app. projecting about as far as the lower branch of the sup. app., its upper margin concave, lowermargin strongly convex. znd, top vieq', f}'avkttitn: Sup. app. as long as 10; its upper branch slender, one-fourtli to one-third us wide at base as long, cylindrical, nearly straight., the equally rounded tip very slightly incurved. Top view, aspersum:
Sup. app. shorter than the side of 10; the upper branch rather broad, its width at
base nearly or quite one-half its length, tapering, the tip slightly expanded inwardly, forming an inner apical tooL11; inner apical margin truncate, oblique; outer apical angle rounded.




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