Elisabeth B. Bryant.
Some Spiders from Acapulco, Mexico.
Psyche 55:55-77, 1948.

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SOME SPIDERS FROM ACAPULCO, MEXICO*
BY ELIZABETH B. BRYANT
Museum of Comparative Zoology
The Museum of Comparative Zoology has recently received two small collections of spiders made at Aca- pulco, by Dr. Harriet E. Frizzell in September 1940, of 4 species and a larger collection of 31 species by Dr. Sarah E. Jones taken in December 1944. Of these 35 species, 8 and an allotype are new. All of these are small and easily overlooked by the casual collector. It is a locality from which the Cambridges had little ma- terial when they wrote the two volumes of the Biologia Centrali-Americana, published in 1889-1905. In a short paper by Dr. W. J. Gertsch in 1907, several species from this locality were included but all were collected at a different season of the year.
The following is a list of the species found. All were collected in December with the exception of the four marked September.
Family (ECOBIIDB
J 2 (Ecobius beatus Gertsch and Davis
Family LYCOSIDB
J $ pullus Sosippus mexicanus Simon, Sept. Family SICARIIDAE
pullus Scytodes c7iampioni F.0.P.-Cambr. ? " liebraica Simon
Family PHOLCIDB
<^ $ P7zysocylus globosus (Tacz.)
Family DICTYNIDB
$ Dictpa sp.
$ " nivea (0.P.-Cambr.)
"Published with a grant from the Museum of Comparative Zoijlogy at Harvard College.
55
Psiche 5555.77 (1948). hup ltpsychc einclub orgtSS155-055 html



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Psyche
[ Jun
Faniily T HERIDIIDZ
$ Latrodect,us rnactans (Fabr.)
$ TJzeridion maculipes spec. nov.
$ " placidum spec. nov.
$ Eperigone ?
Family ARGIOPID-~E
fJ pullus Argiope sp. Sept.
<^ $? Aranea pallidulus (Keys.)
.<j" $ " vesta spec. nov.
$ Eriophora edax (0.P.-Cambr.)
$ Neosconella lineatipes (0.P.-Canibr.)
d' ?
6 i oaxacensis (Keys.) Sept.
31 Tetragnattia alba F.0.P.-Cambr.
<^ ?
i t antillana Simon
<^ 5
i i dentigera F.0.P.-Cambr.
s ?
i i
guat emu1 ensis 0.P.-Cambr.
? pullus Theridiosoma ?
Family THOMISIDX
<^ Misumenops celer (Hentz)
2 8 $ pullus Miszimmops dubhs (Keys.)
Family CLUBIONID^E
$ pullus AnypJima
$ Anyphmella pavida spec. nov.
Family SALTICIDB
J $ Habronattus cambridgei spec. nov.
$ Marpissa melanognatha (Lucas)
Metaphidippis gratis spec. nov.
c?
i i militaris (Hentz)
<^ ?
i i
purus spec. nov.
$ Pl~iale simplicicava (F.0.P.-Cambr.)
å´j Plexippus fawns (Peckham)
Sidusa ?
3 Tomis jonesce spec. nov.




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19481 Bryasnt-Hpiders f row Mexico
Family (ECOBIIDB
Genus (Ecobks Lucas 1845
(Ecobius beatus Gertsch and Davis
Figures 1, 2
GEcobius beatus Gertsch and Davis, 1937, p. 2, figs. 1, 2.
"9 holotype and immature paratypes from Aca- pulco, Mexico, June 17, 1936 (L. I. Davis). " Male. Length, 1.5 mm.
The male is much darker than the holotype but the amount of color evidently depends upon the habitat. The eves are more closely grouped than in the female and are on a low turret. The two black annulae on each joint of the legs are reduced to large lateral dots. The abdominal markings are typical of the genus and are very distinct.
The palpus is large for the size of the spider. There are three conspicuous points all protruding from the cavity, a forked pair in the distal half and a longer point at the base. All are heavily chitinized. Allotvpe 8 Mexico; Acapulco, 19 December 1944 (S. Jones). 38 5$ Mexico ; Acapulco, 18-19 December 1944 (S. Jones).
The male selected as an allotype was found under a rock and has the dark marks on the cephalothorax and the abdomen much more distinct than the specimens found at the base of palm leaves, these are quite pale. Family THERIDIID*~
Genus Theridion Walckenaer 1805
Theridion maculipes spec. nov.
Figure 3
Female. Length, 1.5 mm., ceph. 0.6 mm., abd. 1.1 mni. long, 1.0 mm. wide.
Cephalothorax pale, little darker about the eye area, anterior margin only slightly narrowed, cephalic portion little higher than the thoracic; eyes cover the anterior margin, anterior row recurved, a.m.e. dark, convex, sepa- rated by less than a diameter, a.1.e. smaller than a.m.e., separated from them by a radius of a.m.e., posterior



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row straight, eyes surrounded by red rings, p.m.e. larger than a.m.e., separated by less than a diameter, p.1.e. sepa- rated from p.m.e. by about a diameter of p.l.e., lateral eyes touching, p.1.e. larger than a.1.e. ; quadrangle slightly higher than wide and about the same width in front as behind ; cl ypeus higher than quadrangle ; mandibh~ ver- tical, pale brown; labium dark, fused to the sternum, wider than long ; maxillae pale, more than twice as long as the labium, slightly inclined; sternum white, triangu- lar, about as wide as long, ending in a truncate lobe be- tween the fourth cox% which are separated by fully a di- ameter; abdomen globose, pale, with no definite pattern, a small white inverted V at the base surrounded by black that quickly fades, dorsum with small chitiuized dots that may be the bases of colorless hairs, venter a dull brown with an irregular median white spot; legs relatively short, 1-2-4-3, I1 left missing, white with bro- ken dark rings at the tips of tibiae and metatarsi, no spines but rows of colorless hairs, most conspicuous on the anterior tarsi and metatarsi; epigynum, a pair of heavily cliitinized round opening's just above the fold that touch on the median line with larger and more widely separated sacs beneath the surface just anterior. Holotype ? Mexico; Acapulco, 19 December 1944 (S. Jones).
T71eridion maculipes is very small even for a T71eri- (lion. The most conspicuous characters are the white legs with the black dots and the black inverted V at the base of the abdomen. The openings of the epigynum are very distinct.
Theridion placidurn spec. nov.
Figure 5
Female.
Length, 2.6 mm., cepli. 0.9 mm., abd. 1.6 mm. long, 1.5 mm. wide.
Cephalothorax pale, cephalic portion slightly darker, shining, no hairs; eyes cover the anterior margin, an- terior row slightly recurved, a.m.e. dark, separated by less than a diameter, convex, a.1.e. smaller than a.m.e., separated from them by about a radius of a.iii.tJ., 110s-



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19481 Brymt-Spiders from- Mexico 59
terior row procurved, p.1u.e. larger than a.m.e., separated by less than a diameter, p.1.e. separated from p.1n.e. by a diameter of p.m.e., lateral eyes touching, p.1.e. slightly larger than a.1.e.; qu,adrangle higher than wide, same width in front as behind; clypeus vertical, and higher than quadrangle ; mandibles vertical, pale, cone-shaped ; Ionium pale; maxilla twice as long as labium; abdomen globose, with a narrow median dark stripe with wavering mars-ins outlined with a narrow pale stripe each side. ',
ill the elliarginations a dark spot, three pi& in all, sides a dull tan, entire abdomen with scattered colorless hairs, venter pale ; legs, 1-2-4-3, and I and I1 right legs missing, I pair much the longest, pale, femora and tibiae slightly darker at the tips, rows of colorless hairs; epigynum, a pair of small dark circular sacs, fully two diameters above the fold, separated by a diameter and a half, with small oval openings beneath as figured.
Holotype $ Mexico; Acapulco, 19 December 1944 (S. -
J ones).
Vndoubtedly Theridion placidwn has been confused with Ttieridion spiralis Emerton. The latter is a com- mon species in the northern part of the United States and has been reported by ~~~serling from Venezuela and bv Banks from Teapic, Mexico. Unfortunately a specimen of the Banks identification is not in the mu- s~um collection.
A type specimen of Theridion spira-
Us Emerton from Essex, Massachusetts, which is now before me, has the median stripe on the abdomen pale, heavily outlined by a dark stripe; the epigynum is sepa- rated from the fold by less than a diameter and the lower margin of the area is chitinized and rolled out- wa lad.
Family ARGIOPIDB
Genus Aranea Linnaeus 175.6
Aranea pullidula (Keys.)
l3jwirÌ pu7lidula Keyserling, 1863, p. 124, pi. 4, fig's. 14,15. "$ St. F& de Bogota (N. Granada) ." A~C/I/VI[.<,' pallidulus F.0.P.-Cambridge, 1904, 2, p. 514, pi. 4.9, fig-. 13.




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60 Psyche [June
Keyserling described this species from a single fe- male and states that it is 5.2 rnm. long. Probably because
of poor illumination or magnification, he failed to see the small scape in the middle of the epigynum. F.0.P.- Cambridge had the species from Panama and in the brief description mentions that it is 8.5 mm. long. He also had the type before him. It is a common species at Barro Colorado, Canal Zone, but all specimens seen are about 5.0-5.5 mm. long, so that the size mentioned by Cam- bridge is probably an error. Such a discrepancy in size is of ten misleading.
Aranea vesta spec. nov.
Figures 4, 6
Female.
Length, 5.5 mm., ceph. 2.6 mm., abd. 3.5 mm. long, 3.5 mm. wide.
Cephalothorax dull yellow, paler on cephalic portion, anterior margin about half the greatest width, no thora- cic groove, a pair of long colorless bristles at the end of the cephalic slope; eyes cover the anterior margin, both rows recurved, a.m.e. largest, separated by about a di- ameter with a pair of long colorless bristles between, each bristle from a distinct base, p.m.e. smaller than a.m.e., separated by more than a diameter with one long bristle between, lateral eyes subequal and touching ; quadrangle wider in front and higher than wide; clypeus about a radius of a.m.e. with 6 or 8 long bristles on the margin; mandibles dull yellow, vertical, with scattered long color- less bristles, no boss, fang groove slightly oblique, upper margin with 4 teeth, lower margin with 3 smaller teeth; labium pale brown, wider than long, tip rebordered ; maxi- lla pale, twice as long as labium, sides parallel ; sternum pale, triangular, three-quarters as wide as long, emargin- ate opposite the coxae, pointed at the tip, fourth coxae almost touching; abdomen as wide as long, but with no shoulder angles, a median pale area at the base which rapidly spreads to the width of the abdomen, on the dis- tal half a median shield-shaped dark area, with emar- ginate sides, venter brown with two irregular pale spots anterior to the spinnerets ; spinnerets dark, closely grouped very near the tip; legs, 1-2-4-3, femora dark



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19481 Bryant-&idem from Mexico 61
brown, other joints much paler with faint darker bands near the tips, long black spines, no ventral spines on the femora, III and IV tibiz with dorsal basal spine; epi- gynu~t~, area wider than long, a pale broad median scape with a recurved spoon-shaped tip, base with several short dark bristles, each side of the median scape rather narrow plain areas with outer marquis heavily clCtinized ; a lateral view shows the median scape curved outward, Male.
Length, 3.6 mm., ceph. 2.0 mm. long, 1.5 mm. wide, abd. 2.0. mm. long, 1.6 mm. wide.
Cephalothorax pale brownish-yellow, cephalic por- tion narrowed and rather high, faint thoracic groove on the posterior third, scar of a median bristle midway be- tween eyes and groove ; eyes same as in the female with scars of the two bristles between the a.m.e.; bristles on the margin of the clypeus as in the female; abdomen oval, about two-thirds as wide as long, white with a dark folium on the posterior two-thirds with the emarginate sides as in the female, venter pale brown, with a pair of very faint pale spots anterior to the spinnerets; legs, 1- 2-43, II right missing, pale with vague broad dark bands at the base and tip of tibise, I and II cox= with a small hook, no ventral spines on the femora, spines long and dark, I pair, tibia, ventral, 3-2-2, basal and median pairs very long, lateral, 3-3, dorsal, 1-1-1, metatarsus, ventral, 0, prolateral, 1, retrolateral, 2, 11 pair, patella, at the tip, 1 very long, 1 short, tibia, ventral, 3p short at the base, 2 at the tip, metatarsus, dorsal, 1 at the base, I11 and IV tibiae, dorsal basal spine, IV femur near tip a re- trolateral row of 6 slender spines, III femur, a retrola- teral row of 3 very slender spines ; palpus pale, patella, paracymbium abruptly recurred and the tip broad and brfid; the parts of the palpus best understood from the figure.
Holotype 5 Mexico; Acapulco, 19 December 1944 (S. Jones), Allotype <? Mexico; Acapulco, 18 December 1944 (S. Jones). Paratypes 6 9 8 c? Mexico ; Acapulco, 18 December 1944 (S. Jones).
The generic position of Armea vesta is very uncer- tain. It cannot be placed in the genus Neosconella F.0.P.-Camb. as it lacks ventral spines on the femora



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62 Psyche [June
of both male and female. As the first coxa of the male has a hook, it cannot be placed in the genus Metepek F.0.P.-Camb.
The epigynuni agrees fairly well with the figure of Arafnea detrimentosa (O.P.-Camb.) This species was described from both male and female but F.0.P.-Cam- bridge evidently decided that they were not of the same species and placed the male with Epeira nigrolw- meralis 0.P.-Camb. $ and never placed tlie female originally described under the name detrimentosa. The male of A. detrimentosa has one long bristle 011 the tibia and another from the patella of the palpus and the parts of tlie palpus are quite unlike those of A. vesta. Geims E riophora Simon 1864
Eriophora edax (0.P.-Camb.)
Figure 7
Epeira edax 0.P.-Cambridge, 1863, p. 30. "8 $ Rio
de Janeiro, Brazil. "
The specimen of Eriophora edax from Acapulco ha& a distinct tubercle above the spinnerets and the shoulder angles are slightly developed, both characters unlike the common Eriopfiora balaustina (McCook), found in Florida and Cuba. The abdomen is about 8.0 mm. lo11g and the scape is 4.0 mm. The base is heavily cliitinized and the tip extends beyond the base of the spinnerets. The species has been reported from various parts of Central America and from Puerto Rico.
Genus Neosconella F.O.P.-Cambridge 1903
Neosconella lineatipes (0.P.-Camb.)
Epeira Uneatipes 0.P.-Cambridge, 1889, p. 30, pi. 7, figs. 17, 18.
"$ 8 Guatemala; road between Retalhulen
and Mazatenango, Santa Ana, Guatemala City. " Epim lineatips Keyserling, 1892, p. 190, pi. 9, fig. 141.
Neosconella lineatipes F.O.P.-Cambridge, 1902, p. 476, pi. 45, fig's; 5, 6.
This species is a little smaller than the one described



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19~1 Bryant-Spiders from Mexico 63
by 0.P.-Cambridge but it has the same spots and lines on the anterior pairs of legs, the mandibles and the cephalotliorax. The figure of the epigynum given by 0.P.-Cambridge is very misleading as the scape is turned back and it is not in a normal position. Keyse~ ling has a much better description and the figures of the scape is in the normal position.
$I Mexico; Acapulco, 18 December 1944 (S. Jones). Family CLUBIONIDB
Genus Anyphmella Bryant 1931
Anyphaenella pavida spec. nov.
Figure 12
Female.
Length, 3.0 mm., ceph. 1.0 mm., abd. 2.0 mm. CepJialotJwrax pale, slightly darker about the ante- rior margin, faint darker lateral stripes from the lateral eyes to the posterior margin, moderately convex, thoracic groove faint; eyes, anterior row straight, eyes subequal and equidistant, separated by about a radius, posterior row procurved, slightly longer than the anterior row, separated by more than a diameter ; quadrangle higher than wide and wider behind than in front; clypeus about equals the diameter of a.m.e. ; mandibles brown, vertical, rather long, fang groove transverse, impossible to see the teeth on the fang groove; labium gray, about as long as wide: maxilla pale, not twice as long as labium, sides parallel, not emarginate on the outer margins; sternum pale, nearly as wide as long, convex, widest between I1 coxae, IV cox= separated by less than a diameter; abdo- men pale, shaded with gray in no definite pattern, convex, venter pale, opening of the posterior spiracle one third nearer the spinnerets than to the fold, spinnerets closely grouped ; legs, I right missing, 1-4-2-3, very slender and frail, pale, I pair very long, I tibia; 1.4 mm. long, spines, ventral, 2-2, long and slender, lateral, 2-2, metatarsus, ventral, 2-2; epigynum, area almost twice as wide as long, all the parts beneath the surface deeply colored, a pair of large dark sacs just anterior to the fold, separated by fully a diameter with a notched margin between, in the anterior half and slightly closer together, a pair of



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64 Psyc71 e
[June
oblique openings with cliitinized margins and a darker point at the tip which may be the begimiingoof tubes. Holotype $ Mexico; Acapulco, in palms, IS December 1944 (S. Jones).
Anyp'ha'nella pavida is very delicate and fragile, so 0 oroove
that it is impossible to count the teeth 011 the fang g or the spines on the slender legs but the parts of the epigynum are very distinct. The spider has not ovipo- sited.
Family SALTICIDB
Genus Habronattzis F.0.P.-Cambridge 1901 Habronattus cambridgei spec. nov.
Figures 13, 15, 16
Habronattus viridipes F.0.P.-Cambridge, 1901, p. 224, pi. 21, figs. 4, 5, nec Hentz, 1846; nee Pellenes viri- dipes Peckham, 1885, 1900.
Male.
Length, 4.5 mm., ceph. 2.5 mm. long, 3.5 nm. wide, abd. 2.1 mm. long, 1.6 mm. wide.
Ceplialotliorax dark chestnut brown, with a narrow band of white hairs above the anterior row of eyes and a few scattered white hairs in the ocular area and about the lateral margins, a dark triangle with the apex at the groove that extends to the posterior margin, cephalotho- rax fairly high, widest posterior to the dorsal eyes or between the second pair of legs, thoracic groove short in a circular depression between the dorsal eyes and mid- way in the cephalic plane, thoracic slope quite abrupt; eyes, anterior row recurved so that the upper margins form a straight line, a.m.e. touching, a.1.e. about one- half the diameter of a.m.e. and separated from them by a little more than a line, small eyes about midway be- tween the first and third rows, dorsal eyes subequal with a.1.e.; quadrangle slightly wider behind than in front and about twice as wide as long; clypeus wanting below a.1n.e. and no fringe on the margin ; mandibles dark brown, vertical, long and narrow, fang" groove very short, upper margin with three contiguous teeth, lower margin with one sharp tooth; sternum dark brown, oval, first COX= separated by a diameter, fourth cox% touching; abdomen discolored so that markings are obscured, ven-



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IMSI Bryat~t-Spiders from Mexico 65
ter brown, with lateral darker stripes; legs, 3-4-1-2, Ill pair not modified and no fringes on any leg, brown, with pale rings most conspicuous on the posterior pairs, I pair, femur, prolateral surface slightly flattened and covered with white scales, patella and tibia with many white scales, spines, patella, prolateral, 1, tibia, ventral, 2-2-2, metatarsus, ventral, 2-2, basal pair very long, TI pair with area of white scales on femur smaller than on I pair, III and IV tibiae with dorsal basal spine; palpus not as long as cephalothorax, femur and patella covered dorsally with white scales, tibia and tarsus pale brown, tibia not as long as patella, tibia1 apophysis dark, tarsus &nost circular, bulb about circular, flattend, embolus from a. distinct lobe of the bulb, starting about the middle on the outer side and ending in a very slender tube in a groove at the tip of the cymbium, the second process heavier, from just, below tlic middle of the bulb, with an abrupt turn at the lower margin and ending about opposite the origin on the inner margin.