Nathan Banks.
Notes on Some Hydropsychidæ.
Psyche 43:126-?, 1936.

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Psyche
[December
NOTES ON SOME HYDROPSYCHIDB
Mr. MiIne's third part of his Studies of Trichoptera hav- ing appeared and finishing the series, I offer certain re- marks thereon, particularly on some Hydropsychide. Some- what similar criticisms can be made of various other parts. Mr. Milne has on several occasions credited me with placing a name as a synonym when it was first done by others. Thus under Hydropsyche alternans Walk. he puts H. morosa Bks. (nec. Hag.) as synonym. Now four years be- fore I was born Hagen himself in his Phryganidarum Sy- nopsis Synonomyica, p. 25, 1864, puts his H. moyosa as equal to altemans Walk. McLachlan had indicated it the year before. Both I and Ulmer have simply accepted the dictum of Hagen and McLachlan.
In the type series of H. wzomsa now in the Museum there are five specimens of altermns and three of a Chewrnato- psyche from Red River of the North. These three do not agree we11 with the description, but Milne has taken them to bear the name morosa. Therefore to keep the name morosa as Hagen, Ulmer, and I have kept it for seventy years, I have selected and marked as lectotpye of Hydro- psyche movosa Hagen a male from "St. Lorenz? Canada, Sacken 1859," which is what has been commonly called alternans Walk. The three Cheumatopsyche from Red River are near to some of Hagen's types of H. phalerata. But since Ulmer has figured1 (Selys Monographs fasc. VI, p. 66, 1907) a species of Hydropsyche as Hagen's H. phale- rata (and some of Hagen's types were of this form) I have selected and marked as lectotype Hydropsyche phalerata Hagen a specimen from "Washington, Sacken?" which is the species figured by UImer, This is much smaller than lHis dorsal figure shows apical joint of claspers too long.



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19361 Notes on Some Hydropsychidz 127
the others of the scalaris group, with larger, fewer pale spots, and eyes of male wider apart than the diameter of an eye.
Following Ulmer, Martynov, Mosely, NavAs, and Betten I am separating Cheumatopsyche (that is the Hydropsy- chodes of these authors) on the character used by Betten, the position of certain cross-veins; this appears to be con- stant.
Mosely informs me that the type species of Hydropsy- chodes has no closed median cell, and that Ulmer's figure is wrong; in any event Cheumatopsyche is an older name. Another case is where I am credited with putting H. dubitans as a synonym of N. crepuscular~s, although I had simply followed the statement of McLachlan (Revision Eu- ropean Trichoptera, p. 393, 1878) who had seen the types of both names. As to Walker's Polyoentropus crassicornis, put as a synonym of N. crepuscularis, I examined a type in 1912 and saw that it was a Phylocentropus. Mr. Mosely in- forms me there are four types in the series, two (a male and female) of a Plectroczemia, one female of a Phylocen- tropus9 and one male of a Neureclipsis.
Mosley thinks (and I agree) that the name should be kept for the Plectrocnemia? and I select as lectotype of Polyce%- tropus crassicornis Walk. a male in the type series which is a Plectrocnemia,; all types are from Georgia. Milne lumps under H. scaZaris about all the forms that have the same general structure of the penis, ignoring fur- ther differences. In the male type of H. scalaris the apical joint of the clasper has a distinct hook at the end; the eyes of the male, although larger than in the female, are wider apart than the width of an eye. Betten, in sinking my H. hagewi? has called attention to Hagen's statement that the eyes of the male scala~?ds were larger than the female. This is but a half-truth, for Hagen also says after the descrip- ton that he has a male with still larger eyes, and adds "IS it different ??'.
I have selected and marked as lectotype of Hydropsyche scalaris Hagen a male from "St. Lorenz, Canada, Sacken." H. hageni is smaller and darker than H. scalaris, the male



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128 Psyche [December
eyes are less than the width of an eye apart. It is common
in southern localities where the typical scahris is not found. The last joint of the clasper is shorter and broader than in scalaris.
H. incornmoda Hagen from Georgia is also put as a synonym by Milne. The superior plate is more nearly Fig. I. Male genitalia of Hydropsyche. 1, H. separluta, clasper and plenis; 2? H, reciprocu, clasper and penis, and superior plate; 3, H. incommo~da, clasper; 4? H. scalak, clasper; 5, H. sepamta, superior plfate; 6, H. dep~avata, superior plate; 7, H. scalar%, superior plate, penis below
and above; 8, H. incommoda, superior
plate, penis, two specimens; 9, H. phaleruta, superior plate? penis; 10, H. hagmi, clasper? apical joint.
conical, with only a short incision, the apical joint of the clasper has the hook less distinct. The eyes of male are wide apart, furth,er apart than in H. scalaris. It occurs also in Louisiana.




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19361 Notes on Some hydro psych id^ 129
There are minor differences in these four forms, a11 of which, scalaris hageni, incommoda, phale~a~ta, I consider distinct species, judging by the characters used in the sepa- ration of European species.
Mr. Milne has put at least two (perhaps three) forms under Hydropsyche depravata Hagen.
H. depravata was
described from a female from Georgia; a male from Georgia (same handwriting) appears to be the same species and Milne makes it the neoallotype. This has the apical part of the clasper slender, curved, and scarcely nar- rowed toward tip; the superior plate has a deep, triangular emargination.
In New England, Eastern New York, and Pennsylvania there is a fairly common species of the same group as H. depravat~. It is as large as H. scalaris, darker and with more numerous and smaller spots, and in the male the eyes widely separated. The female is readily separated from H. scalaris by having the mid basitarsus dark and scarcely swollen (in scalaris extremely broad and very pale).
The apical part of the clasper is hardly as long as in H. depravata and plainly tapering toward the tip; the superior plate is deeply, acutely indented, the lobe each side being narrowed near the tip and pointed. This is probably H. reciproca Walk., and I shall so consider it. The type of Walker's species is a female and has lost the middle legs. H. depravata is probably the same. From the western part o$ New York and further west is a form similar in size and markings.
The superior plate
has only a rather shallow, rounded emargination, and the apical part of the clasper is plainly somewhat clavate to- toward tip. This is certainly a different form from the Eastern species and I have named it
Hydropsyche sepanta sp. nov.
The fore wings are much spotted with gray, few in ex- treme apical part; the penis, seen from side, is rounded at tip, slightly swollen and almost pointed above (in recip-



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130 Psyche [December
roca not at all pointed above) ; the superior plate has a me- dian rounded emargination, which when not shriveled, is as wide as deep; the male eyes wide apart. Apical part of claspers slightly enlarged at tip. The female has the mid basitamus dark and but little swollen. The fore wing is 11 mm. long.
Type (and paratypes) from Westfield, N. Y., June and July 1905 (Woglum coll.) . Others are from Saskatchewan, Scudder.
Polycentropus confusm Hagen was described from three specimens, two from Trenton Falls N. Y., and a female from Washington, D. C. One, a female, from Trenton Falls, N. Y. has the discal cell in the hind wings closed and is a Plectrocnemia, and is so treated by Betten, necessitating a change in PlectrocrLemia confma, Mosely. This specimen when 1 first saw the collection and until lately was the first specimen in the row. But the other specimen from Trenton Falls, N. Y. is a male, a true Polycentropus. In 1914 (Can: Ent.) I figured the genitalia of this species as PolycerLtro- pus confusw; I have therefore selected this Trenton Falls male as lectotype of Polycentropus confwus Hagen and so marked it. Thus Mosely's Plectrocnemia will not need renaming.
The selection of these lectotypes will result in keeping several of the Hagen names as they have been treated and figured, which, I believe, is better than to make the changes which have recently been put forward, and in each case the specific names is retained in the genus in which Hagen de- scribed it.




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