F. B. Shaw.
Notes on the Mycetophilidæ with Descriptions of New Species.
Psyche 42:84-90, 1935.

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Psyche
[June
NOTES ON THE MYCETOPHILIDB WITH
DESCRIPTIONS OF NEW SPECIES
Cornell University, Ithaca, N. Y.
In working over material preparatory to writing the chapter on the Mycetophilidse to be published in the Diptera of Connecticut, a few observations were made that are be- lived to be best published before that work appears. Not a great deal of work has been done in this country on this group of insects since the appearance of the Genera In- sectorum fascicle by Johannsen in 1909. Later, in a series of bulletins published by the Maine Agricultural Experiment Station, the same author gave keys to all of the known spe- cies occuring in this country.
Since that time Edwards of the British Museum has done the most outstanding work in the group.
He revised the
family in 1924 and as it is the belief of the writer that the paper of Edwards is not easily accessible to the average worker on Diptera, it is considered to be worthwhile to give a summary of Edwards' revision, giving the arrangement of the group according to Johanrisen and comparing it with the classification of Edwards.
In Johannsen's revision of the group, nine subfamilies were recognized. According to Edwards' grouping there are ten subfamilies but they are not equivalent in all cases to those of Johannsen which bear the same name. Since it is the belief of the writer that there are many of the same or closely related species occuring both in Europe and in Amer- ica, it is felt that a classification should be adopted which is uniform. Consequently, the revision of Edwards is being followed in this paper. The following table lists only those genera found in North America :
Pu&f 42:84-91 (1935). hup Ytpsychu einclub nrg/42/42-084 html



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19351 Notes on Mycetophilidse 85
Subfamilies and Genera
Subfamilies and Genera
(Johannsen) (Edwards)
Subfamily Bolitophilinse Subfamily Bolitophilinas Bolitophila Bolitophila
Hesperinus
Subfamily Pachyneurinse
Subfamily Mycetobinse Subfamily Ditomyiinse Mycetobia
Palseoplatyura
Ditomyia Ditomyia
Symmerus Symmerus
-
Subfamily Diadocidiinse Subfamily Diadocidiinse Diadocidia
Diadocidia
Subfamily Ceroplatinse Subfamily Ceroplatinas Asindulum Asindulum
Ceroplatus Ceroplatus
Cerotelion
Hesperodes Hesperodes
Apemon Apemon
Platyura Platyura
Palseoplatyura
Subfamily Macrocerinee Subfamily Macrocerinse Macrocera
Macrocera
Subfamily Sciophilinse
Monoclona
Eudicrana
Tetragoneura
Sciophila
Paratinia
Polylepta
Empalia
Dziedzickia
Neoempheria
Mycomya
Diomonus
Subfamily Sciophilinse
Monoclona
Eudicrana
Tetragoneura (in part)
Sciophila (in part)
Paratinia
Polylepta (in part)
Synapha
Dziedzickia (in part)
Neoempheria
M ycomyia
Allocotocera
Leptomorphus
Neuratelia
Syntemna
Phthinia
Megalopelma




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Psyche
Sciophila
Hadroneura
Sciophila
Acnemia
Azana
Speolepta
Ccelosia
Hadroneura
Gnoriste
Boletina
Rondaniella
Leia
Pnyxia
Docosia
Megophthalmida
Ectrepesthoneura
Subfamily Mycetophilinse Subfamily Mycetophilinse Gnoriste
Probolseus
Acnemia
Azana
Rondaniella
Odontopoda
Leptomorphus
Allocotocera
Boletina
Leia
Phthinia
Ccelosia
Syntemna
Megophthalmidia
Docosia
Anatella
Trichonta
Cordyla
Brachypeza
Rhymosia
Allodia
Phronia
Telmaphilus
Exechia
Dynatosoma
Opistholoba
Epicypta
M~c~othera
Mycetophila
Anatella
Trichonta
Cordyla
Brachypeza
Rhymosia
Allodia
Phronia
Exechia
Dynatosoma
Epicypta
Mycetophila (in part)




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Notes on Mycetophilidse
Sceptonia
Zygomyia
Delopsis
Sceptonia
Z ygomyia
Delopsis
Subfamily Sciarinse Subfamily Sciarinse
Eugnoriste
Manota
Pn yxia
Trichosia Trichosia
Zygoneura Zygoneura
Metangela
Phorodonta Phorodonta
Rhynchosciara
Sciara Sciara
The chief difference between the classification of Johann- sen and that of Edwards will be noted in the limits of the two subfamilies Sciophilinse and Mycetophilinse. Johannsen, 1911, recognized that there were two quite distinct groups in the Mycetophilinse and gave a classification of these two series based on the arrangement of setulse on the wing. Edwards, 1924, become convinced that a more natural grouping would be obtained by placing the genera included in Series I of the Mycetophilinse of Johannsen in the Scio- philinse.
Since some of the genera have been split or united with other genera since the date of Johannsen's work, I believe it best to mention the more important changes that have oc- curred.
1. The genus Hesperinus was removed by Johannsen to the Bibionidse.
2. Pachyneura is now placed in a separate family re- lated to the Anisopodidse.
3. Mycetobia is placed with the Anisopodidae by some authors or in a distinct family the Mycetobiidse. 4. Palseop~latyura is placed with the Ceroplatinse by Edwards.
5. Cerotelion is considered to be a subgenus of Cero- platus.
6. Diomonus is united with the genus Leptomorphus.



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88 Psyche [June
7. Megalopelma includes one species of the genus Phthinia and probably will include some of the North Am- erican species of Sciophila.
8. Speolepta was erected for Polylepta leptogaster of Winnertz.
9. Ectrepesthoneura includes those species of Tetragon- eura having the cubital fork near the base of the wing. Ac- cording to Garrett, this genus is not valid but as the writer has not seen the evidence to support this view, he does not care to make any decision.
10. Probolseus is united with the genus Lygistorhina of the subfamily Lygistorhininse.
11. Manota is placed in a distinct subfamily the Mano- tinse.
12. Telmaphilus is united with the genus Phronia. 13. Opistholoba is united with the genus Mycetophila. 14. Mycothera is united with the genus Mycetophila. 15. Pnyxia is considered to belong with the Sciophilinse according to Edwards.
In the course of the investigation a few specimens were encountered that are worthy of a more detailed discussion. Two of these were found to have been originally recorded from Greenland and from Europe. Spechens were sent to Edwards for an examination and he confirmed the identifi- cation. The writer wishes to acknowledge the assistance given to him by Doctor Edwards at this point. Boletina groenlandica Staeg.
Naturh. Tidsskr 1 : 356
(PI. VI, Fig. 1.)
A specimen of this insect was taken by Dr. C. P. Alexan- der from Hermit Lake, Mt. Washington, New Hampshire. There is no figure of this species published in America so that a figure of the clasper (fig. 1) is given. AIIodia ornaticollis Meigan. Syst. Bescher 1 : 269. (PI. VI, Fig. 2.)
This is the first record of the capture of this European species in America. Since there is no figure published in



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193.51 Notes on Mycetophilidss 89
this country of this species, the hypopygium (fig. 2) is pic- tured. Taken in Ithaca, N. Y. Nov. 1934. Exechia aviculata sp. n.
(PI. VI, Fig. 3.)
Male. Length 4 mm. Head brown, palpi and antennae light brown, basal two joints of antennae yellowish. Thorax brown, humeri not tinged with yellow. Abdomen brown, the last three segments somewhat darker. Coxse and femora with a brownish cast, tibia and tarsi yellow. Prothoracic basitarsus about one fourth longer than tibia, tibia swollen at the apex.
Hypopygium (fig. 3) light brown resembling somewhat that of Exechia janmrii Lunds.
Type locality,
Ithaca, New York. Type in my collection, paratype in Cornell University collection.
Exechia pollex sp. n.
(PI. VI, Fig. 4.)
Male. Length 3.5 mm. Head dark brown, palpi and an- tennse brown, basal two segments of the antennae light. Thorax dark brown, humeri with a light yellowish tinge. Abdomen concolorous, dark brown. Coxae and femora yel- low, tibia and tarsi somewhat darker. Prothoracic basitar- sus and tibia subequal in length. Hypopygium (fig. 4) re- sembles that of Exechia, frigida but differs in the details of the claspers. Type locality, Ithaca, New York. Type in my collection.
Rhymosia triangularis sp. n.
(PI. VI. Fig. 5.)
Male and female. Length 4-4% mm . Head pale brown, palpi and basal two joints of antennae yellow, remainder of antennae brown. Mesonotum brown, lateral margins some- what lighter.
Coxae and femora yellow, femora slightly darkened at apices. Tibia and tarsi brownish yellow. Pro- thoracic basitarsus about a third longer than the tibia. Dor- sum of first and sixth abdominal segm,ents brown. The re-
maining segments with subtriangular brown spots with the apex of each segment as the base of the triangle. Hypo- pygium (fig. 5) yellow, resembling that of Rhymosa inflata but one of the arms of the forceps is much more slender. Type locality, South Bethlehem, New York. Type in my collection, paratype in Cornell University collection.



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90 Psyche [June
Psyche, 1935
VOL. 42, PLATE VI.
-.
Shaw-Mycetophilidae




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19351 Notes on Mycetophilidse,
EXPLANATION OF PLATE VI.
Figure 1.
Clasper of Boletina grcenlandica.
Figure 2.
Hypopygium of Allodia ornaticollis.
Figure 3.
Hypopygium of Exechia aviculata.
Figure 4.
Hypopygium of Exechia pollex.
Figure 5.
Lateral view of clasper of Rhymoskx triangularis. A; arm of clasper.
Drawings made by D. B. Creager.
A NEW NAME FOR LITHOMYRMEX CARP.
(HYMENOPTERA)
In my revision of the fossil ants (Bull. Mus. Comp. Zool., 70 (1) : p. 36), which was published in January, 1930, I es- tablished the myrmicine genus Lithomyrmex for two species, rugosus and striatus, from the Miocene shales of Florissant, Colorado. Unfortunately, the same generic name was used by Dr. J. Clark a few months earlier for a recent species of ant, glauerti, from West Australia (Journ. Roy. Soc. W. Australia, 14: p. 36, 1929). I therefore propose the new name Eulithomyrmex for Lithomyrmex Carp. (nee Litho- myrmex Clark, 1929), to include the two extinct species from the Florissant shales-F. M. Carpenter.



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