P. J. Darlington, Jr.
New West Indian Carabidae, with a list of the Cuban Species.
Psyche 41:66-130, 1934.

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66 Psyche [June
NEW WEST INDIAN CARABID^E, WITH A LIST
OF THE CUBAN SPECIES1
Museum of Comparative Zoology, Cambridge, Mass. The purpose of the present paper is two-fold: first, to describe a number of new Carabidse which have come to hand during several years of sporadic work on the West Indian fauna, and second, to give a complete list of the species now known from Cuba. Eventually I plan to publish a revision of all the Carabidse of the West Indies. How- ever, this will have to await the acquisition of much more material than is now available, and I think in the mean- time a' Cuban list (which adds very little to the length of this paper) will be useful not only to collectors on the island, but to any coleopterists in the United States who feel a responsibility for knowing something about the Cuban fauna.
The order of genera in the list is approximately that of the Junk Catalogue. The Cuban species are numbered con- secutively; species from other islands are interpolated without numbers. Of the 134 species here listed from Cuba (about twice the number previously known) I have seen
Cuban specimens of 128. The 6 which I have not seen are noted in the list, The synonymy which I have cited is practically limited to names used for actual Cuban speci- mens by earlier writers, of whom the chief have been the following :
1.
Jaequelin-Duval, 1857, in Ramon de la Sagra's His- toire physique, politique et naturelle de 1'Ile de Cuba (French Ed.), Animaux Articules [Vol. 71, pp. 6-24 (Carabidse) .
'Publication aided by a grant from the Museum of Comparative Zoology.
including as a supplement a new Masoreus (Aephnidius) by A. J. Mutchler.




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19341 . New West Indian Carabidse 67
2.
Chevrolat, 1863, Ann. Soc. Ent. France (4) 3, pp. 186- 199.
3. Gundlach, 1891, Contribucih ii la Entomologia Cubana, Vol. 3, part 5, pp. 12-33 (sometimes cited as of An. Acad. Cien. Habana).
4. Leng & Mutchler, 1914, Bull. American Mus. Nat. Hist. 33, pp. 393-397.
5. Leng & Mutchler, 1917, Ibid. 37, pp. 194-195. The material I have examined includes the collections of the Museum of Comparative Zoology (containing Cuban specimens collected by me in 1926 and 1929, on trips made possible by grants from the Atkins Foundation of Harvard University), the United States National Museum, the American Museum of Natural History, and the Philadelphia Academy of Sciences (Poey material from Cuba) ; collec- tions submitted by Prof. Stuart T. Danforth (Puerto Rico), Mr. S. C. Bruner of the Estacih Experimental Agronomics and Mr. M. L. Jaume (Cuba) ; and smaller lots of specimens from other sources. My thanks are due to all of these per- sons and to the curators of the museums mentioned. I have also to acknowledge assistance given me by Mr. Ren6 Oberthiir, Mr. K. G. Blair, Dr. R. Jeannel, Dr. Joseph Bequsert, Mr. A. d'orchymont, Mr. W. S. Blatchley, Mr. A. J. Mutchler, Mr. M. Banninger, Dr. S. Breuning, and the late Prof. H. F. Wickham, all of whom have either com- pared specimens with types inaccessible to me, or identified material in special genera, or helped me in other ways. 1.
Calosoma (s. s.) splendidurn Dej.
2.
Calosoma ( Callistriga) a. alternans (Fab. ) 3. Pachyteles gyllenhali (Dej.)
pallida (Chev.)
4. Scarites alternans Chd.
5. Scarites subterraneus Fab. (varities) 6. Clivina dentipes De j.
Clivina addita n. sp.
Of average form for Clivina of dentipes group, slightly depressed ; black, appendages piceous. Head with clypeus



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68 Psyche [June
bi-emarginate each side, but less deeply so than in dentipes; front with longitudinal and transverse lines as in dentipes, with small median puncture, otherwise impunctate ; an- tennss not reaching to basal angles of prothorax. Pro- thorax by measurement very slightly wider than long; moderately convex; posterior margin basal; posterior an- gles not conspicuous; disk not punctate, with usual im- pressed lines and also with fine, irregular, well separated transverse lines. Elytra not margined at base, rather deeply striate, striae moderately punctate; eighth stria not continued above humerus ; third stria five-punctate. Front femur with small, rounded-obtuse tooth on lower posterior edge near apex; front tibia tridentate externally above terminal digit, finely bicanaliculate on anterior face near apex, inconspicuously toothed near middle of posterior face ; front trochanters not angulately prominent; middle tibia spurred externally near apex; paronychium short or absent in unique type, but probably long in fresh specimens. Lower surface without unusual punctuation ; last ventral with inner pair of punctures twice as far apart as distance between them and outer punctures. Length 8 mm.
Holotype (Museum of Comparative Zoology no. 19489) from Mayaguez, Puerto Rico, F. Mara, sent by S. T. Dan- forth ; unique.
In Putzeys' revision of Clivina (Ann. Soc. Ent. Belgique 10, 1866) this species would belong in group 24 (dentipes etc.), but it will not fit in any of the six sections of that group. It is excluded from the first four sections either by absence of a basal elytral margin or by the small obtuse femoral tooth; from the fifth, by the impunctate abdomen and only slightly and broadly emarginate epistoma; and from the sixth, by the short antennae. It is at once separ- able from de-es, which it somewhat resembles, by the reduced femoral tooth.
7. Clivina cub= n. sp.
Slender, parallel, slightly depressed; piceous black, ap- pendages and parts of lower surface rufescent. Head with clypeus deeply bi-emarginate each side ; front shining, al-



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19341 New West Indian Ca'rabidse 69
most impunctatle except for a punctiform impression at middle; antennae reaching about to basal angles of pro- thorax. Prothorax depressed, almost square, not narrowed in front; anterior angles minutely prominent, posterior angles finely denticulate ; impressed lines on disk normal for dentipes group; surface of pronotum very finely punctate. Elytra with striae moderately impressed and punctate ; not margined at base between first and sixth intervals; small tubercle at base of first stria; third stria with five setigerous punctures; sixth and seventh intervals finely carinate at base, sixth joining elytral margin in front of humeral angle. Front femur with a short, obtuse tooth (hardly more than a pronounced sinuation) below near apex; front tibia lightly sulcate above, tridentate externally above terminal digit, with only a very minute tooth near middle of posterior face; front trochanters not angulately prominent; middle tibia with spur on outer side near apex; paronychium about as long as claws. Pro,-, meso-, and metathorax punctate at sides below; abdomen finely but distinctly punctate, espec- ially at sides and on apical segment; prosternum about as in dentipes; last ventral with inner pair of setae separated by about one and a half times the distance between them and outer setae. Length 5%-6 mm.; width -+- 134 mm. Holotype (Museum of Comparative Zoology no. 19490) and 10 paratypes from Soledad (near Cienfuegos) Cuba, June and Oct. 31, washed from gravel banks of the Arimao River; 6 paratypes from Cayamas,l Cuba, May 25-June 8, E. A. Schwarz (United States National Museum) ; 1 para- type from Baragua, Camaguey, Cuba, June 5, L. D. Chris- tenson, at light (U. S. N. M.) .
This species belongs in group 24 (dentipes group) in Put- zeys' revision (I. c.). Within this group it falls in' section 5 or 6, resembling only C. latimana Putz. and C. punctiven- tris Putz., both of South America, in ventral punctua- tion. From latimom, CU~SB differs in having the epistoma 'The Cayamas from which Schwarz's specimens came is in south- western Santa Clara, near Yaguararnas
(Cf. Proc. Ent. Soc. Wash-
ington 5, 1903, p. 287).




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70 Psyche [June
not more emarginate than usual at middle; from punctiven- tris, in having shorter antennae. A comparison of speci- mens would probably show other differences. From Clivina dentipes and C. addita, the only previously described West Indian species of its group, cubas is distinguished at once by its smaller size and more extensively punctate lower surface. 8. Clivina limbipennis J.-Duval
simplex Chlev.
9. Clivina insularis (J.-Duv.)
10. Clivina bipustulata (Fab.)
bipustulu (Chev.)
11. Clivina biguttata Putz.
bisignatu Chev. etc., not bisignata Putz. 12. Dyschirius subiaevis Putz.
13. Dyschirius erythrocerus Lec.
14. Oxydrepanus rufus (Putz.)
brevicarinutus (Putz. )
15. Ardistomus nitidipennis n. sp.
Of average form for Ardistomus; black, not bronzed, very shining; appendages and a pair of large, rounded, scarcely oblique subapical spots rufous. Head with clypeus almost evenly truncate; vertex not impressed. Prothorax almost orbicular; lateral margins reaching base ; disk with usual impressions. Elytra very shining, not at all aluta- ceous ; striae tnoderately impressed, impunctate, entire, sec- ond reaching nearly to base ; third interval five-punctate on outer edge (constant in all specimens). Last ventral with two punctures each side near apex. Front tibia bidentate externally above apical digit; front tarsi rather widely di- lated. Length 5%-6% mm.
Holotype (Museum of Comparative Zoology no. 19491 ) and 12 paratypes from Soledad (near Cienfuegos) Cuba, June, Oct. 21, Nov. 7 & 9; some taken in debris after heavy floods, others secured by treading .down vegetation on the edge of a tiny brook in woods.




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19341 New W& Indian Carabidse 71
This is the first red spotted Ardistomus known from the West Indies.
It belongs in group 1, section 1 of Putzeys' revision (Ann. Soc. Ent. Belgique 10, 1866, 200-). It is close to A. convexa Putz., of which we have a broken speci- men from Mexico, but differs in being slightly less convex and in having the elytra shining, not alutaceous as in con- vexa. The Cuban species is also brighter, with larger ely- tral spots. From A. obliqwta Putz. of the eastern United States, including Florida, the Cuban species differs in being smaller, with different outline of prothorax, and with fewer setigerous punctures on the third elytral interval. 16. Ardistomus elongatulus Putz.
17. Ardistomus cyaneolimbatus Chev.
gundlachi Putz. MS., Gundlach, etc.
18. Aspidoglossa vulnerata Putz.
comma Putz.
19. Schizogenius arimao n. sp.
Subparallel, slightly depressed ; rufous (immature) or black, appendages and lower surface always more or less rufescent. Head with mentum toothed ; vertex s'even-sul- cate, central carinae nearly parallel, only slightly converging anteriorly. Prothorax by measurement slightly wider than long; disk impunctate, slightly wrinkled transversely, with a single longitudinal sulcus each side of the median sulcus. Elytra rather deeply striate, striae punctate basally, not apically; third and fifth intervals each with about seven, seventh with about four setigerous punctures (exact num- ber slightly variable). Front tibia as usual in genus. Male with three setigerous punctures placed triangularly each side of last ventral; female with two setae each side, near margin. Length 33/^% mm.
Holotype s (Museum of Comparative Zoology no. 19492) and 24 paratypes from Soledad (near Cienfuegos) Cuba, June, Oct. 19, Dec. 1 & 3; all washed from low gravel banks of the Arimao River. One paratype from Cumanayagua, Santa Clara, Cuba, June 10, I?. de Zayas (in Coll. Estacih E. Agronomica, Cuba).




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72 Psyche [June
This is the first known West Indian Schizogenius. It is close to S. tristriatlis Putz. of Mexico and Central America (several specimens examined) but is smaller and more con- vex than typical tristrwtus (described as 5 mm. long) and much smaller than variety longipennis Putz. (described as larger than tristriatus) . Bates, in "Biologia," gives the size of his specimens of tristriatus as from 1% to 2% lines, which would include specimens as small as the Cuban ones. However, the identity of the various Central American forms referred to tristriatus seems to me very questionable, and since the Cuban form, if it really is tristriatus at all, is fairly constant and smaller than either of the described con- tinental forms, it seems best to treat it as a distinct species at least temporarily. It can hardly prove to be less than a good subspecies. In Leconte's table of North American Schizogenius (Bull. Brooklyn Ent. Soc. 2, 1879, p. 34) arimao runs to lineolatus or saZZei, but is smaller and more convex than either.
The following key will help to define the numerous new species of Bembidiini which are described in the following pages. It includes all previously known West Indian species except Tachys pictwatus Putz. (possibly a member of the vittiger group) and T. piceolus Laf. (possibly near corrus- cus), which I have not been able to identify in the material I have seen. It also excludes the completely blind Petro- charts eggersi Ehlers of St. Thomas, which I do not know. 1. Outer apical angle of front tibia normal, rounded; size ......
2%-6 mm. .......................................... (Bembidion) .2 Outer apical angle of front tibia obliquely truncate or truncato-emarginate ; size 3 mm. or (usually) less ...... 6 2. Dorsal elytral punctures practically on third stria ; color plain dark rufous ; length 4 mm. ................................ Bembidion jamaicense n. sp.
Dorsal elytral punctures near middle of third interval; elytra more or less variegated .......................................... .3



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19341 New West Indian Carabida 73
...........
3. Front of head not at all alutaceous ; 2%-3y3 mm.. Bembidion darlingtoni Mutchler.
...........................
Front of head alutaceous ; size larger.. .4
4. Elytral striae apically deleted or only lightly impressed. Bembidion viridicolle (Laf.) .
.....
Elytral striae entire, about same depth throughout.. .5
5. Prothorax with sides broadly but slightly sinuate before basal angles ; latter only slightly prominent.. .................... Bembidion fastidiosum (Laf.) .
Prothorax with sides strongly and conspicuously sinuate ...................
before basal angles ; latter rather prominent Bembidion sparsum Bates.
6. Two setae over each eye, but anterior sometimes shorter .................................................................. than posterior.. .7
One seta over each eye; head sunk in thorax nearly to eyes .................................................... (Micratopus) ....... .27 7. Eyes moderate or large; antennae sometimes monili- form, usually not. .............................................................. .8 Eyes much reduced, small or minute; antennae monili- form ...................................................... (Limnastis) ....... .28 8. Antennae moniliform, middle joints as wide as long.. ... 9 Antennae more slender.. ................................................... .I0 9. Side margins of thorax broad, channeled, translucent; color castaneous with base and apex of elytra pale; size .......................................................................... under 2 mm.
Tachys (Tachyta) flavicauda autumnalis Bates. Side margins of prothorax narrower, not translucent; color dark piceous, elytra each with a subapical red spot ; size -+- 2V2 mm.. .................................................................. Tachys (Tachyta) hispaniolae n. sp.
10. Elytra each with six coarsely punctate strise beside the marginal series of punctures; size -+-2-2s mm ............. Pericompsus blomdulus Schaum.
Elytra with not more than five coarsely punctate striae beside marginal series, or striae not coarsely punctate .I1



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74 Psyche 1 June
11. Posterior dorsal puncture of elytron (on outer edge of third interval, as a rule) much in front of apical re- curved stria ; body very convex .................................... .12 Posterior dorsal puncture of elytron on or within hooked tip of apical recurved stria; body usually more de- pressed ............................................................................... .13 12. Elytra each with five coarsely punctate striae or rows of .......................................
punctures ; mentum bi-perforate,
Tachys (subgenus?) immaculatus (Bates).
Elytra each with only one or two striae, which are im- punctate ; mentum not perforated ..................................... Tachys (Tachyura) xanthopus Dej.
13. Posterior elytral puncture directly upon hooked tip of apical recurved stria ; surface not iridescent.. ............. .14 Posterior elytral puncture within, not on, hooked tip of apical recurved stria; surface usually more or less iri- ............................................................................... descent .15,
14. Basal angles of prothorax obtuse. ....................................... Tachys bradycellinus Hayw.
Basal angles of prothorax right. .Tachys occultator Casley (and) Tachys ensenadae Mutchler.
15. Male with two joints of front tarsus dilated .............. .16 Male with only basal joint of front tarsus dilated ...... .20 16. Antennae very slender (middle joints about three times as long as wide; tip of sixth reaching about to basal an- ...
gle of prothorax) ; eyes moderate; length ~3 mm. Tachys albipes Lee.
Antennae stouter and short~er; other characters vari- able ..................................................................................... .17 17. Eyes abruptly prominent, forming right angles with sides of head behind them; basal angles of prothorax obtuse but distinct; length just over 2% mm. ................... Tachys abruptus n. sp.
Eyes less prominent, forming obtuse angles with sides ...........
of head behind them; other characters variablce 18




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19341 New West Indian Carabidse 75
18. Basal angles of prothorax very obtuse, almost rounded except for minute subprominent sinuation of margin; ..................................................................... size &21/^ mm..
Tachys putzeysi F. & S.
Basal angles of prothorax obtuse but rather distinct; .................................................................. size 2-214 mm.. .19
19. Prothorax transverse, head relatively; narrow (PI. 1, fig. 1) ....................................... .Tachys dominicanus n. sp. Prothorax less transverse, head broader. ........................... Tachys sp. (Guadeloupe & Montserrat.)
20. Surface of elytra with very fine, short, sparse pubes- .................................................... cence Tachys striax n. sp.
......................................
Surface of elytra not pubescent
21
21. Posterior angles of prothorax accurately and minutely formed ; color testaceous with head scarcely darker .. .22 Posterior angles of prothorax finely blunted or bluntly subdenticulate; color castaneous, or partly testaceous with head much darker.. ......................... ..'. ..................... .25 22. Form unusually slender, parallel.. ................................. .23 ..................................................................... Form broad. .24
23. Base of prothorax oblique at sides; eyes very prominent. Tachys flax n. sp.
Prothorax squarely truncate at base; eyes Less promi- nent .................................................... Tachys pumzlus Dej. 24. Median line of prothorax continued as a groove behind posterior transverse impression; first joint male front tarsus very wide ; size -+-2% mm.. ....................................... Tachys cubax n. sp.
Median line of prothorax not continued behind posterior transverse impression; first joint male front tarsus rela- tively much narrower; size just under 2 mm. .................... Tachys paulax n. sp.
25. Sides of prothorax sinuate before basal angles, which would be almost right except are slightly blunted; head and prothorax relatively narrow; size *2l,4 mm ........... Tachys scitulus Lee.




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76 Psyche [ June
Sides of prothorax scarcely or not sinuate before the basal angles, which are more obtuse; other characters ............................................................................. variable .26
26. Color usually castaneous or darker; head and prothorax relatively narrower ; antennae more slendler (middle joints more than twice as long as wide) ; first joint male front tarsus wider and with outer apical angle more strongly produced ; size -h 2% mm.. ................................... Tachys corruscus Lec.
Color testaceous with head much darker; head and pro- thorax relatively widler ; antennae stouter (middle joints less than twice as long as wide) ; first joint male tarsus smaller; size 2 mm. or slightly less ................................... Tachys vorax Lec.
27. Head moderate ; prothorax not especially large ; size about 1% mm.. ...................... Micratopus insularis n. sp. Head very small; prothorax relatively very large ; size +2 mm. (PI. 5, fig. 3) ......... Micratopus parviceps n. sp. -
28. Form slender; head relatively small; length 1%-1% mm. (PI. 5, fig. 2.) ........................ Limnastis americanus n. sp. Form less slender ; head relatively large ; length barely over 1 mm.. ................................... Limnastis capito Bates. Bembidion (Peryphus) jamaicense n. sp.
Moderately slender, convex, upper surface faintly aluta- ceous but rather shining; dark rufous, appendages testa- ceous. Head rather long ; eyes prominent ; antennae slender ; front with inconspicuous median puncture; frontal sulci moderate, parallel; mentum with rounded-triangular tooth at middle. Prothorax subcordate, about one-third wider than long (by measurement) ; basal angles acute but not much more than right, strongly carinate; disk with trans- verse impressions moderate, longitudinal line strong. Elytra