C. T. Brues.
Ænictomyia, a New Myrmeeophilous Phorid from the Philippines.
Psyche 37:163-166, 1930.

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19301 New Myrrnecophilous Phorid
SNICTOMYIA, A NEW MYRMECOPHZLOUS
PHORZD FROM THE PHILIPPINES
Among the extensive collections of ants obtained by Dr. J. W. Chapman on the island of Negros in the Philippines are several specimens of a most remarkable species of Phoridae which he found living with ants of the genus Xnictus. Dr. Chapman has very kindly placed these speci- mens at my disposal with the suggestion that I describe the species, which proves to be a wholly unknown form, which must form the type of a new subgenus. In the very recent classification of this family by Schmitz, the new senictophile falls in the genus Diploneura, which Schmitz has divided into three subgenera, Diploneura sensu stricto, Tristoechia and Dohrniphora, but it differs from all of these by a very complete development of the dorsal macrochsetae on the thorax as well as by the excessive length of the bristles on the head and wings. The following tabular arrangement will indicate the characteristics that serve to distinguish the several sub- genera :
1.
Hind tibiae with two dorsal (extensor) lines formed of closely spaced and imbricated bristles, and be- tween these with a series of minute well separated bristles ; mesopleurse bare above.. .............................. .2 Hind tibiae with only a single row of bristles form- ing a line of the dorsal surface, mesopleurae hairy above ............................................................................... .3 2. Fourth longitudinal vein sinuous, the base curved upwards and the apex downwards Diploneura, s. str. Fourth longitudinal vein straight, not distinctly sinu- ous, nor curved at base of apex. ................ Tristcechia



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164 Psyche [ June
3.
Mesonotum completely bristled; with a median row of four macrochsetse; four achrostichals, three or four dorsocentrals and six marginal scutellar bris- tles; the macrochsetse all as long or longer than the mesonotum; base of third vein bearing one very long bristle ... ... .. . . . . . .. .. . ... .. ... .. . .. . ... Bnictomyia, subgen. nov. Mesonotum minutely hairy, without macrochsetse ex- cept near lateral and posterior margins ; base of third vein bare or with one or two very short bristles Dohrinphora
Aenictomyia subgen. nov.
Although clearly related to Dohrniphora, the type of this subgenus exhibits several striking characters. The bris-
tling of the entire body is unusually long as well as the cilia on the costal vein. The very complete and conspicuous chsetotaxy of the mesonotum is very unusual, as in prac- tically all Phoridse the median and anterior macrochsetse are completely suppressed, although those along the pos- terior and lateral margins are generally more or less well developed. The front is completely bristled ; the proboscis geniculate and strongly chitinized; eyes hairy; palpi strongly bristled; cheeks each with three very strong mac- rochsetse ; propleura hairy, with a large macrochseta ; meso- pleura hairy above; front tibia with a series of four small macrochsetse on dorsal surface; middle tibia with a strong basal pair; hind tibiae without macrochsetse before apex; fourth wing-vein curved ; seventh not distinct. Type : Diploneura (Bnictomyia) chaprnani sp. nov. Diploneura (Aenictomyia) chapmani sp. nov., Fig. 1. Female : Length 1.2-1.5 mm.
Yellowish brown, the middle portion of the front, the mesonotum more or less, the tergites of the second and third abdominal segments and the apical one darker brown, the legs lighter, except the hind pair; wings hyaline with the heavy veins light brown. Front wider than long, the



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19301 New Myrmecophilous Phorid 165
surface smooth, impunctate ; the upper margin sharp and forming a slightly raised rim except at the sides; two closely approximated supra-antenna1 bristles directed up- wards and backwards; lower transverse frontal row con- sisting of four equidistant bristles, the lateral ones well removed from the margin of the eye; middle row further Fig. 1.
Diploneura (^Emctomyia) chapmmi sp. nov. from the lower one than from the upper, of four equi- distant bristles; upper or ocellar row close to the occipital margin ; postocular cilia strong, especially above ; cheeks each with three very large and strong bristles ; bristles of palpi rather strong and densely placed. Proboscis beyond the basal angle about as long as the head-height, gently curved downwards to the tip. Antennae small, oval; with a moderately long, weakly pubescent arista. Mesonoturn smooth, highly polished, without any microscopic hairs be-



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166 Psyche [June
tween the large macrochsetse; six scutellar bristles, the median ones much further from one another than the others so that there is a group of three on each side, the middle one of which is much longer than either the lateral or median one. Propleura sparsely hairy, with four small bristles at the base of the coxa and a large, erect one at the upper angle; mesopleura sparsely hairy on the upper half. Front tibia with a series of four moderately long bristles on the outer (extensor) edge, one at the basal third, one at middle and two between middle and apex. Middle tibia with a pair of long bristles near base (Fig. 1, c) and two short ones externally at tip. Hind tibia with- out long bristles. Abdomen very weakly chitinized ; with a distinct large dorsal plate on the second and a small trans- verse plate on the second segment. Wings rather small, costa extending beyond the middle (Fig. 1, b), its cilia very long; third vein forked well before apex; base of third vein with one very long bristle near base; seventh vein not distinct.
Described from three specimens taken by Dr. J. W. Chapman with ^Enictus martini Forel, at his summer camp in the mountains near Dumaguete, Negros Oriental, Phi- lippine Islands. Dr. Chapman tells me that the Phorids ran along in the foray during the migration of the colony, making faster progress than the ants.




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