James M. Carpenter.
The identity of Odynerus scudderi Cameron and O? bradleyi Cameron (Hymenoptera: Vespidae: Eumeninae).
Psyche 94:77-80, 1987.

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THE IDENTITY OF ODYNERUS SCUDDERZ CAMERON AND 0? BRADLEY1 CAMERON
(HYMENOPTERA: VESPIDAE: EUMENINAE)*
BY JAMES M. CARPENTER
Museum of Comparative Zoology, Harvard University, Cambridge, MA 02138
Bohart (1965) treated the identity of 100 names of North Ameri- can Eumeninae described by Peter Cameron from 1905 to 1912. This paper established the synonymy of all but four of these names, and so solved many problems in the nomenclature of nearctic Eumeninae. During a recent visit to the British Museum, I studied the type specimens of two of the unrecognized species, and in the present work establish their synonymy.
Euodynerus scudderi (Cameron, 1908) NEW COMBINATION Odynerw scudderi Cameron, 1908: 196. Q; locality not stated (British Museum). Odynerus discogaster Bequaert, 1939: 65.9; "Mt. Diablo, Contra Costa Co., 1,500 to 2,000 ft., CALIFORNIA"(Museum of Comparative Zoology). NEW SYNONYMY. Bohart (1965) stated in regard to this species: "Type location unknown; pseudotype Q, Chimney Gulch, Colorado, BMNH #18.541, does not agree with original description." In fact, it agrees with the original description perfectly, and I regard it as the true type. It bears a handwritten label "Chimney Gulch Col. 6/ 17/98". The writing is identical to that on material of other Cameron species collected by Oslar and specifically referred to in the same paper. Cameron's description did omit various details which he commonly included in other descriptions, such as the coloration of the legs, and the label data. But the details which he did report match the speci- men, and are sufficiently precise to dispel1 doubt. In particular, the features of the clypeus are convincing. Cameron stated (1908: p. 197): "Clypeus distinctly longer than wide; in its centre is a black mark extending from top to bottom, the top square, from there it becomes gradually widened to the middle, then gradually narrowed to near the apex, which extends in a clearly separated angle; the 'Manuscript received by the editor August 15, 1986. Parhi Ìö4:77-8 (1087). hup Ytpsycht einclub orgÌö4f94-077.htni



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78 Psyche [vo~. 94
sides of the mark are irregular, the apex is transverse." This is distinctive, as coloration goes. Further, he mentioned (p. 196): a 'mark over the antennae, broader than long, transverse above, gradually roundly narrowed below, ending there in a nipple-like point", which is also distinguishing. Finally, he also mentioned (p. 197): "The head, thorax and base of the abdomen are densely covered with long, fuscous black hair." This is a specific character of Euodynerus discogaster (Bequaert, 1939), which now becomes a junior synonym. I have compared the holotype of this latter species with the holotype of fiodynerus scudderi, and scudderi differs primarily by the absence of the discal spots on metasomal tergum 11. Among the specimens of discogaster in the MCZ is a female labelled "Webster Colorado July" which also lacks these spots and confirms that discogaster is variable in this regard. Pterochilus [!] decorus Cresson, 1879: xvii. $5 (Bohart, 1940, states that the female is the holotype); "Nevada" (Academy of Natural Sciences). Odynerus ? bradleyi Cameron, 1909: 81. "$" (= 8); "Durango, Colorado" (British Museum). Senior secondary homonym in Pterocheilus of Pterocheilus bradleyi Bohart, 1950, NEW SYNONYMY
Bohart did not find the type of this species. I chanced upon a specimen labelled in Cameron's hand "Odynerus ? bradleyi Cam." It lacks a BM type number label, but was probably not assigned one because "type9' was not written on the determination label, contrary to Cameron's usual practice. The specimen is labelled with a printed label reading "Oslar. Durango Col.;" beneath this is a handwritten label reading "Durango Col 5/29/99;" a label bearing the number "5 197;" the printed label "P. Cameron Coll. 19 14-1 10;" and the aforementioned determination label. The handwritten labels are in the same hand as other specimens stated to be from Oslar. The type has the head glued In place, the forelegs and propleura broken off, the midlegs (the left coxa remains), and the right hind tibia and tarsus also broken off. One of the fore coxae and femur, the right midleg and the right hind tibia and tarsus are glued to the first label. The specimen is otherwise intact, and is easily recognizable as a species of Pterocheilus. The specimen matches the (for Cameron) extended description quite well, both in coloration and structure.



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19871 Carpenter- Odynerus scudderi
Distinctive details are the distribution of rufous color on the meta- soma, the (Cameron, 1909: p. 81) "Clypeus not much longer than wide, the top dilated broadly in the middle, the apex with a shallow incision," the "Antennae longish, stout, dilated towards the apex," the pronotal "keel," and the metasoma! shape (I "cup-shaped," I1 "as wide as long and narrowed at the base;" actually it is one and one third times as wide as long). However, Cameron stated that the specimen is a female, but it is a male. The description of the antenna! apex leaves no doubt that Cameron was dealing with a male and erred in stating the sex. He stated: "the apical joints marked with black at the apex, the last rounded above, flat below, roundly narrowed at the apex."This is a common feature of mates in the subgenus Megapterocheilus, to which this species belongs. Pterocheilus bradleyi Cameron, 1909 is a junior subjective synonym of Pierocheilus decorus Cresson, 1879.
Pterocheilus chesteri NEW NAME
Pterocheitus (Megaptetocheilus) bradleyi Bohart , 1950: 195.8; "Nogaies, Arizona" [Corneli University). Junior secondary homonym in Pterocheilus of Odynerus ? bradleyi Cameron, 1909
Unfortunately, the discovery of the identity of 0. bradleyi Cameron creates a case of secondary homonymy in Pterocheilus. P. bradleyi Cameron, although a junior synonym, is an available name (Article 10(g) of the International Code of Zoological Nomencla- ture), and as such is a senior homonym of P, bradleyi Bohart, 1950. This latter species, described from Arizona in the same subgenus, must be rejected (Art. 59(a)), and replaced by a new name (Art. 60(c)), as it has no junior synornyms. I herewith propose the name Pterocheilus chesteri for bradleyi Bohart. The original name was proposed by Bohart as a patronym for J. Chester Bradley, and that intent is maintained by the replacement name. Work in the British Museum was supported by a Milton Fund grant to the author. I thank M. C. Day for assistance during my stay, and for valuable comments on the initial draft of this paper.



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Psyche
[Vol. 94
BEQUAERT, J.
1939.
Odynerus annectens de Saussure and related species, with additional notes on Odynerus hidalgo de Saussure (Hymenoptera, Vespidae). Ann. Entomol. SOC. Amer. 32: 58-69.
BOHART, R. M.
1940.
A revision of the North American species of Pterocheilus and notes on related genera (Hymenoptera, Vespidae). Ann. Entomol. Soc. Amer. 33: 162-208.
1950.
North American additions to the genus Pterocheilus (Hymenoptera, Vespidae). Proc. Entomol. Soc. Wash. 63: 195-202. 1965.
Synonymy of North American Odynerini described by Peter Cameron (Hymenoptera: Eumenidae). Pan-Pacif. Ent. 41: 107-1 13. CAMERON, P.
1908.
A contribution to the knowledge of the Odyneridae of the Southwest of the United States. Trans. Amer. Entomol. Soc. 34: 195-228. 1909.
On some Diplopteryga from the Southwest of North America. Pomona J. Ent. 1: 78-85.
CRESSON, E. T.
1879. Trans. Amer. Entomol. SOC. Proc. 7: xvii.



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