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PSYCHE

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Wm. T. M. Forbes.
Two Wasp-Guests from Puerto Rico.
Psyche 40:89-93, 1933.

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PSYCHE
VOL. XL SEPTEMBER, 1933 No. 3
TWO WASP-GUESTS FROM PUERTO RICO
(MICROLEPIDOPTERA) .
Cornell University, Ithaca, N. Y.
The two following species were reared by Mr. Francisco Seh, Jr., from nests of Polistes, along with the well-known Setomorpha insectella, and two Pyralids, which are to be included in Dr. Schaus's revision of that family. One of these new species which is accompanied by the pupa has also been found in Cuba, making a description perhaps more advisable.
Taeniodictys, new genus
Head with smooth flat occiput, heavily tufted vertex, and smooth, strongly retreating front. Ocelli invisible, appar- ently absent; eyes large, labial palpi moderate, obliquely upturned or drooping (loosely held), clothed with irregular bristly scales, but without true bristles; maxillary palpi of folded type, the visible part short, drooping on each side of tongue, which is naked. Antennae with scape broad, con- cave and bristly-scaled, covering over half of eye but hardly a true eyecap, without pecten; shaft 5/6, smoothly scaled with one whorl to a segment. Body nearly cylindrical, the cox= not flattened against the body; mid-tibise smooth- scaled, normal, hind tibiae with spurs at 2/5, clothed above with bristly hairs, which are long except before the upper Pu&e 40339.93 (1933). hup Ytpsychu einclub orgt4W40-089 html



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Psyche f September
spurs; hind metatarsus also clothed above with some bristly hairs. Fore wing (fig". 1) linear, rather caudate; smooth and glossy; Sc short, Ri absent, Ra and Rs free, arising from the upper side of the cell, which is represented by a ridge on the under side, there being no trace of a true vein; R4 stalked with M2, Mi lost; Ms normal, from same point of origin as ptalka of R4-Ms; Cu simple, a true vein all the way, exceedingly closely parallel to the ridge that represents R, so that the cell is linear; A short, sinuous, simple; 1st A represented by a slight fold only; hind wing under 1/2; fringe over 6; costa with sinuousity and spe- cialized scales quite close to base, the vein Sc fading out only a little beyond it; Cu simple and short; the remaining veins represented only by ridges on the under side, which appear to represent Rs and three medians, the two upper connate; no cell.
This genus would belong to the OinophiIid~e, considering the well developed palpi (labial and maxillary) and semi- scavenger habits. By Meyrick's grouping it would be in- cluded in the Lyonetiidse, whose typical members have lost the mouth-parts, and have smooth heads, and have larvae feeding on living plants. The genus appears distinct, but in the complete absence of any attempt at a classification of the group it is difficult to be sure. I have not come across any genus in the group which shows the exceedingly narrow cell, and most of the few genera which have lost Ri are Old World forms of quite different appearance. Tsaniodictys sericeUa, new species
Glossy ash gray, nearly immaculate.
Top of head and
thorax rather paler, with Iuteous tint, gradually shading into luteous on lower part of tuft and under aide of head, antenm pale pearl gray above, the scape more luteous ; fore wing shading gradually into deeper gray toward apex and luteous white toward inner margin, the basal half of dorsal fringe markedly shaded with luteous.
Hind wing clay
color, the dorsal fringe also heavily shaded with brighter luteous. Legs dirty white, shaded on outer sides with light gray. Abdomen pale gray, the under side toward apex and anal tuft paler and yellower. 6-7 mm.




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19331 Wasp Guests from Puerto Rico 91
Lares, Puerto Rico, Dec. 1931, holotype and several para- types reared from nests of PolIStes crinttus by Fr. Sein, Jr., with a much larger number of the following species. Holo- type in Cornell University collectio,n, type No. 1240. Antipolistes, new genus
Similar to Tinea. Head densely hairy all over, the hair above the antennae tending to form two tufts, as usual; eyes very small, separated by more than 3 times their width; ocelli invisible; antennas over 2/3, simple, scape simple, somewhat roughly scaled, without pecten or eyecap, shaft with one whorl of long scales to a segment, smooth toward base, somewhat rough outwardly; labial palpi moderate, drooping, second segment bristled, third rather shorter, fusiform ; maxillary palpi invisible, and tongue absent ; hind tibia with upper spurs between 1/3 and 1/4, the outer upper spur very long, reaching nearly to tip of tibia; loosely hairy above and with some loose hair below; tarsus not modified. Wings lanceolate (figs. 2 $,3 Q ) ; fore wing with Ri and Cui lost, and one other vein (apparently Mi) ; the three apical veins stalked, but not always the same way; no accessory cell; Rg running to costa, Ra arising at 5/6, Cua a little further out but hardly at angle.
1st A absent, 2nd A
simple.
Hind wing broad lanceolate, 2/3 fore wing, the costa a little sinuate beyond middle; subcosta ending be- yond the sinuation but far before apex, R simple, free, Mi lost, Ms very weak and free, Ms longer and free; Cu a much stronger vein, plainly forked at apex. No cell or anal veins. Fringe 3.
There are a few weak aculese at the base of the cell of the fore wing, besides the usual patch on the inner margin. This genus is the most reduced of the Tineidse known to me, the vein formula being 9-5; I judge it is a reduction of Achanodes, but the only genera that seem to have reached a similar stage of reduction in venation are the Indian Brachydoxa and Asyndetaula, which show by their short antennae and smooth hind tibiae that they do not belong to the true Tinea group. Meyrick's Clepticodes hexaleuca (Exot. Micr., iv, 324) must be similar, but if correctly



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92 Psyche
[September
placed by Meyrick has a much more complete structure :- with maxillary palpi folded, Rs apparently free, and all three medials of hind wing preserved, but Cui lost. Antipolistea anthracella, new species
Coal black. Head lighter, wood brown to umber brown according to the light, becoming darker on under side of tuft and about mouth. Antennae and palpi fuswus gray; legs pearl gray, the segments barred with dark gray, leav- ing pale bands at the joints, all very difficult to see dearly on account of the &sa of the scaling. Fore wing coal black, with four whitish spots, on the costa near middle and at 5/6, and in the fold at 1/3 and rather below the fold at 2/3; the spots variable in size, and tending to be rel- atively smaller in the female. Under side mouse gray. Female much like male, but noticeably larger, and appar- ently darker. s 4 1/2 mm. e nearly 6 mm. Larea, Puerto Rim, holotype male and numerous sped- mens of both sexea reared by Fr. Sein, Jr., from nests of Po Ws enkhs, in Dec. 1931 ; also several paratypes from Guabairo, Central Soledad, Cienfuegos, Cuba, reared from two nests of Polistes cubensts Lep. by Richard Dow. Hob
type in Cornell University collection, type no. 1241, para- types in Cornell University collection, Museum of Compara- tive ZoQlogy and National Museum.
The larva appears to be unknown. Pupa (from Sein) similar to Tinea peZIWneUa as figured by Miss Moaher (Bull. 111. State Lab. Nat. Hist., xii, pi. 20, figs. 30, 311, sutures not clearly seen, prothorax depressed dorsally, maxillae shorter, nearly obsolete, but maxillary palpi large and transverse ; antennae barely longer than fore wings ; angula- twins of terminal segment ventral rather than lateral- The small round eyes are also more completely visible below the antennae. Abdomen dorsally with first three segments unarmed; fifth and sixth with anterior rows of fine spinules; 7th and 8th also with coarser rows near the mid-segments, the latter rather sparse; ninth segment with two subdorsal clusters of about three spinules each and last with small pointed subdorsal cones,




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Psyche 1933
Vol. 40, Plate 4.
Forbes-Microlepidopterous Wasp-Guests




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Volume 40 table of contents